@import url('https://fonts.googleapis.com/css2?family=Inter:wght@400;500;600&family=JetBrains+Mono:wght@400&display=swap');

*, *::before, *::after { box-sizing: border-box; margin: 0; padding: 0; }

:root {
  --canvas: #f7f7f4;
  --canvas-soft: #fafaf7;
  --surface-card: #ffffff;
  --surface-strong: #e6e5e0;
  --primary: #f54e00;
  --primary-active: #d04200;
  --ink: #26251e;
  --body: #5a5852;
  --body-strong: #26251e;
  --muted: #807d72;
  --muted-soft: #a09c92;
  --on-primary: #ffffff;
  --hairline: #e6e5e0;
  --hairline-soft: #efeee8;
  --hairline-strong: #cfcdc4;
  --semantic-success: #1f8a65;
  --semantic-error: #cf2d56;
  --radius-xs: 4px;
  --radius-sm: 6px;
  --radius-md: 8px;
  --radius-lg: 12px;
  --radius-xl: 16px;
  --radius-pill: 9999px;
}

html { scroll-behavior: smooth; }

body {
  background: var(--canvas);
  color: var(--body);
  font-family: 'Inter', system-ui, 'Helvetica Neue', Helvetica, Arial, sans-serif;
  font-size: 16px;
  font-weight: 400;
  line-height: 1.5;
  -webkit-font-smoothing: antialiased;
}

a { color: var(--ink); text-decoration: none; }
a:hover { color: var(--primary); }

img { max-width: 100%; height: auto; display: block; }

.container {
  max-width: 1200px;
  margin: 0 auto;
  padding: 0 24px;
}

/* TOP NAV */
.top-nav {
  background: var(--canvas);
  border-bottom: 1px solid var(--hairline);
  height: 64px;
  position: sticky;
  top: 0;
  z-index: 100;
}
.top-nav .container {
  display: flex;
  align-items: center;
  justify-content: space-between;
  height: 64px;
}
.nav-brand {
  font-size: 18px;
  font-weight: 600;
  color: var(--ink);
  letter-spacing: -0.01em;
  line-height: 1;
}
.nav-brand span { color: var(--primary); }
.nav-links {
  display: flex;
  align-items: center;
  gap: 32px;
  list-style: none;
}
.nav-links a {
  font-size: 14px;
  font-weight: 500;
  color: var(--body);
  line-height: 1.4;
  transition: color 0.15s;
}
.nav-links a:hover { color: var(--ink); }
.nav-links a.active { color: var(--ink); }
.nav-hamburger {
  display: none;
  background: none;
  border: none;
  cursor: pointer;
  padding: 8px;
  flex-direction: column;
  gap: 5px;
}
.nav-hamburger span {
  display: block;
  width: 22px;
  height: 2px;
  background: var(--ink);
  border-radius: 2px;
  transition: all 0.2s;
}

/* HERO */
.hero-band {
  padding: 80px 0;
  background: var(--canvas);
}
.hero-band h1 {
  font-size: 72px;
  font-weight: 400;
  line-height: 1.1;
  letter-spacing: -2.16px;
  color: var(--ink);
  max-width: 820px;
  margin-bottom: 24px;
}
.hero-band .hero-sub {
  font-size: 16px;
  color: var(--body);
  max-width: 600px;
  line-height: 1.5;
  margin-bottom: 32px;
}
.hero-meta {
  font-size: 13px;
  color: var(--muted);
  margin-bottom: 48px;
}
.hero-image {
  width: 100%;
  max-height: 480px;
  object-fit: cover;
  border-radius: var(--radius-lg);
  border: 1px solid var(--hairline);
  margin-top: 48px;
}

/* BADGE */
.badge-pill {
  display: inline-block;
  background: var(--surface-strong);
  color: var(--ink);
  font-size: 11px;
  font-weight: 600;
  letter-spacing: 0.88px;
  text-transform: uppercase;
  border-radius: var(--radius-pill);
  padding: 4px 10px;
  margin-bottom: 20px;
}

/* SECTIONS */
.section {
  padding: 80px 0;
}
.section-alt {
  background: var(--canvas-soft);
  border-top: 1px solid var(--hairline-soft);
  border-bottom: 1px solid var(--hairline-soft);
}
.section-title {
  font-size: 36px;
  font-weight: 400;
  line-height: 1.2;
  letter-spacing: -0.72px;
  color: var(--ink);
  margin-bottom: 16px;
}
.section-sub {
  font-size: 16px;
  color: var(--body);
  max-width: 600px;
  margin-bottom: 48px;
}
.section-label {
  font-size: 11px;
  font-weight: 600;
  letter-spacing: 0.88px;
  text-transform: uppercase;
  color: var(--muted);
  margin-bottom: 12px;
}

/* CARDS */
.cards-grid {
  display: grid;
  gap: 24px;
}
.cards-grid-3 { grid-template-columns: repeat(3, 1fr); }
.cards-grid-2 { grid-template-columns: repeat(2, 1fr); }

.feature-card {
  background: var(--surface-card);
  border: 1px solid var(--hairline);
  border-radius: var(--radius-lg);
  padding: 24px;
}
.feature-card h3 {
  font-size: 18px;
  font-weight: 600;
  color: var(--ink);
  line-height: 1.4;
  margin-bottom: 10px;
}
.feature-card p {
  font-size: 14px;
  color: var(--body);
  line-height: 1.5;
}
.feature-card .card-icon {
  font-size: 24px;
  margin-bottom: 16px;
}

/* ARTICLE CARDS */
.article-card {
  background: var(--surface-card);
  border: 1px solid var(--hairline);
  border-radius: var(--radius-lg);
  overflow: hidden;
  transition: border-color 0.15s;
}
.article-card:hover { border-color: var(--hairline-strong); }
.article-card img {
  width: 100%;
  height: 200px;
  object-fit: cover;
}
.article-card-body { padding: 24px; }
.article-card-body .article-meta {
  font-size: 11px;
  font-weight: 600;
  letter-spacing: 0.88px;
  text-transform: uppercase;
  color: var(--muted);
  margin-bottom: 10px;
}
.article-card-body h3 {
  font-size: 20px;
  font-weight: 400;
  letter-spacing: -0.11px;
  color: var(--ink);
  line-height: 1.3;
  margin-bottom: 10px;
}
.article-card-body p {
  font-size: 14px;
  color: var(--body);
  line-height: 1.5;
  margin-bottom: 16px;
}
.article-card-body .read-link {
  font-size: 14px;
  font-weight: 500;
  color: var(--ink);
  border-bottom: 1px solid var(--hairline-strong);
  padding-bottom: 1px;
  transition: color 0.15s, border-color 0.15s;
}
.article-card-body .read-link:hover { color: var(--primary); border-color: var(--primary); }

/* SPECIES TABLE */
.species-table {
  width: 100%;
  border-collapse: collapse;
  font-size: 14px;
}
.species-table th {
  text-align: left;
  font-size: 11px;
  font-weight: 600;
  letter-spacing: 0.88px;
  text-transform: uppercase;
  color: var(--muted);
  padding: 12px 16px;
  border-bottom: 1px solid var(--hairline);
}
.species-table td {
  padding: 12px 16px;
  border-bottom: 1px solid var(--hairline-soft);
  color: var(--body);
  vertical-align: top;
}
.species-table td:first-child { color: var(--ink); font-weight: 500; font-style: italic; }
.species-table tr:last-child td { border-bottom: none; }
.table-wrap {
  background: var(--surface-card);
  border: 1px solid var(--hairline);
  border-radius: var(--radius-lg);
  overflow: hidden;
}

/* ARTICLE PAGE */
.article-header { padding: 64px 0 48px; }
.article-header h1 {
  font-size: 48px;
  font-weight: 400;
  letter-spacing: -1.44px;
  line-height: 1.15;
  color: var(--ink);
  max-width: 760px;
  margin-bottom: 20px;
}
.article-header .article-meta {
  font-size: 13px;
  color: var(--muted);
  display: flex;
  gap: 20px;
  flex-wrap: wrap;
}
.article-body {
  max-width: 760px;
}
.article-body h2 {
  font-size: 26px;
  font-weight: 400;
  letter-spacing: -0.325px;
  color: var(--ink);
  margin: 48px 0 16px;
  line-height: 1.25;
}
.article-body h3 {
  font-size: 20px;
  font-weight: 600;
  color: var(--ink);
  margin: 32px 0 12px;
  line-height: 1.3;
}
.article-body p {
  font-size: 16px;
  color: var(--body);
  line-height: 1.6;
  margin-bottom: 20px;
}
.article-body ul, .article-body ol {
  margin: 0 0 20px 24px;
  font-size: 16px;
  color: var(--body);
  line-height: 1.6;
}
.article-body li { margin-bottom: 6px; }
.article-body img {
  width: 100%;
  border-radius: var(--radius-lg);
  border: 1px solid var(--hairline);
  margin: 32px 0;
}
.article-body .img-caption {
  font-size: 13px;
  color: var(--muted);
  margin-top: -24px;
  margin-bottom: 32px;
}
.article-body .info-box {
  background: var(--surface-card);
  border: 1px solid var(--hairline);
  border-left: 3px solid var(--ink);
  border-radius: 0 var(--radius-md) var(--radius-md) 0;
  padding: 20px 24px;
  margin: 32px 0;
  font-size: 15px;
  color: var(--body);
  line-height: 1.6;
}
.article-body .ref-list {
  font-size: 13px;
  color: var(--muted);
  margin-top: 48px;
  padding-top: 24px;
  border-top: 1px solid var(--hairline);
}
.article-body .ref-list a { color: var(--muted); text-decoration: underline; }
.article-body .ref-list a:hover { color: var(--body); }
.article-layout {
  display: grid;
  grid-template-columns: 1fr 300px;
  gap: 64px;
  align-items: start;
  padding-bottom: 80px;
}
.article-sidebar { position: sticky; top: 80px; }
.sidebar-card {
  background: var(--surface-card);
  border: 1px solid var(--hairline);
  border-radius: var(--radius-lg);
  padding: 24px;
  margin-bottom: 20px;
}
.sidebar-card h4 {
  font-size: 11px;
  font-weight: 600;
  letter-spacing: 0.88px;
  text-transform: uppercase;
  color: var(--muted);
  margin-bottom: 16px;
}
.sidebar-card ul { list-style: none; }
.sidebar-card ul li { margin-bottom: 8px; }
.sidebar-card ul li a {
  font-size: 14px;
  color: var(--body);
  line-height: 1.4;
}
.sidebar-card ul li a:hover { color: var(--ink); }

/* FORM */
.contact-form {
  background: var(--surface-card);
  border: 1px solid var(--hairline);
  border-radius: var(--radius-lg);
  padding: 40px;
  max-width: 600px;
}
.form-group { margin-bottom: 20px; }
.form-group label {
  display: block;
  font-size: 14px;
  font-weight: 500;
  color: var(--ink);
  margin-bottom: 8px;
}
.form-group input,
.form-group textarea {
  width: 100%;
  background: var(--surface-card);
  color: var(--ink);
  border: 1px solid var(--hairline-strong);
  border-radius: var(--radius-md);
  padding: 12px 16px;
  font-size: 16px;
  font-family: inherit;
  line-height: 1.5;
  height: 44px;
  transition: border-color 0.15s;
  outline: none;
}
.form-group textarea { height: auto; min-height: 120px; resize: vertical; }
.form-group input:focus,
.form-group textarea:focus { border-color: var(--ink); }
.form-group input::placeholder,
.form-group textarea::placeholder { color: var(--muted-soft); }
.btn-primary {
  background: var(--primary);
  color: var(--on-primary);
  font-size: 14px;
  font-weight: 500;
  line-height: 1;
  padding: 10px 18px;
  height: 40px;
  border-radius: var(--radius-md);
  border: none;
  cursor: pointer;
  transition: background 0.15s;
  display: inline-flex;
  align-items: center;
  gap: 8px;
}
.btn-primary:hover { background: var(--primary-active); }
.btn-primary:disabled { opacity: 0.6; cursor: not-allowed; }
.form-message {
  display: none;
  margin-top: 16px;
  padding: 12px 16px;
  border-radius: var(--radius-md);
  font-size: 14px;
  font-weight: 500;
}
.form-message.success {
  background: #d1f0e6;
  color: var(--semantic-success);
  border: 1px solid #a3e0cc;
}

/* DIVIDER */
.divider { height: 1px; background: var(--hairline); margin: 0; }

/* PAGE HEADER */
.page-header {
  padding: 64px 0 48px;
  border-bottom: 1px solid var(--hairline);
}
.page-header h1 {
  font-size: 48px;
  font-weight: 400;
  letter-spacing: -1.44px;
  line-height: 1.15;
  color: var(--ink);
  margin-bottom: 16px;
}
.page-header p {
  font-size: 16px;
  color: var(--body);
  max-width: 600px;
}
.page-body {
  padding: 64px 0 80px;
  max-width: 760px;
}
.page-body h2 {
  font-size: 26px;
  font-weight: 400;
  letter-spacing: -0.325px;
  color: var(--ink);
  margin: 40px 0 14px;
}
.page-body h3 {
  font-size: 20px;
  font-weight: 600;
  color: var(--ink);
  margin: 28px 0 10px;
}
.page-body p { font-size: 16px; color: var(--body); line-height: 1.6; margin-bottom: 16px; }
.page-body ul, .page-body ol { margin: 0 0 16px 24px; font-size: 16px; color: var(--body); line-height: 1.6; }
.page-body li { margin-bottom: 6px; }

/* FOOTER */
.footer {
  background: var(--canvas);
  border-top: 1px solid var(--hairline);
  padding: 64px 0 48px;
}
.footer-grid {
  display: grid;
  grid-template-columns: 2fr 1fr 1fr 1fr 1fr;
  gap: 40px;
  margin-bottom: 48px;
}
.footer-brand p {
  font-size: 14px;
  color: var(--body);
  line-height: 1.6;
  margin-top: 12px;
  max-width: 280px;
}
.footer-col h5 {
  font-size: 11px;
  font-weight: 600;
  letter-spacing: 0.88px;
  text-transform: uppercase;
  color: var(--muted);
  margin-bottom: 16px;
}
.footer-col ul { list-style: none; }
.footer-col ul li { margin-bottom: 10px; }
.footer-col ul li a {
  font-size: 14px;
  color: var(--body);
  transition: color 0.15s;
}
.footer-col ul li a:hover { color: var(--ink); }
.footer-bottom {
  border-top: 1px solid var(--hairline);
  padding-top: 24px;
  display: flex;
  justify-content: space-between;
  align-items: center;
  flex-wrap: wrap;
  gap: 12px;
}
.footer-bottom p { font-size: 13px; color: var(--muted); }
.footer-bottom a { font-size: 13px; color: var(--muted); }
.footer-bottom a:hover { color: var(--body); }

/* COOKIE BANNER */
.cookie-banner {
  position: fixed;
  bottom: 0;
  left: 0;
  right: 0;
  background: var(--ink);
  color: var(--canvas);
  padding: 20px 0;
  z-index: 200;
  transform: translateY(100%);
  transition: transform 0.3s ease;
}
.cookie-banner.visible { transform: translateY(0); }
.cookie-banner .container {
  display: flex;
  align-items: center;
  justify-content: space-between;
  gap: 24px;
  flex-wrap: wrap;
}
.cookie-banner p { font-size: 14px; color: var(--canvas-soft); line-height: 1.5; flex: 1; min-width: 260px; }
.cookie-banner a { color: var(--canvas); text-decoration: underline; }
.cookie-actions { display: flex; gap: 12px; flex-shrink: 0; }
.btn-cookie-accept {
  background: var(--primary);
  color: var(--on-primary);
  font-size: 13px;
  font-weight: 500;
  padding: 8px 16px;
  border-radius: var(--radius-md);
  border: none;
  cursor: pointer;
  transition: background 0.15s;
}
.btn-cookie-accept:hover { background: var(--primary-active); }
.btn-cookie-reject {
  background: transparent;
  color: var(--canvas);
  font-size: 13px;
  font-weight: 500;
  padding: 8px 16px;
  border-radius: var(--radius-md);
  border: 1px solid rgba(255,255,255,0.3);
  cursor: pointer;
  transition: border-color 0.15s;
}
.btn-cookie-reject:hover { border-color: rgba(255,255,255,0.6); }

/* BREADCRUMB */
.breadcrumb {
  display: flex;
  gap: 8px;
  align-items: center;
  list-style: none;
  font-size: 13px;
  color: var(--muted);
  padding: 20px 0;
}
.breadcrumb li + li::before { content: '/'; margin-right: 8px; color: var(--hairline-strong); }
.breadcrumb a { color: var(--muted); }
.breadcrumb a:hover { color: var(--body); }

/* DISCLAIMER */
.disclaimer {
  background: var(--canvas-soft);
  border: 1px solid var(--hairline);
  border-radius: var(--radius-md);
  padding: 16px 20px;
  font-size: 13px;
  color: var(--muted);
  line-height: 1.5;
  margin: 32px 0;
}

/* UPDATED DATE */
.updated-date {
  font-size: 13px;
  color: var(--muted);
  margin-bottom: 4px;
}

/* RESPONSIVE */
@media (max-width: 1024px) {
  .hero-band h1 { font-size: 56px; letter-spacing: -1.5px; }
  .cards-grid-3 { grid-template-columns: repeat(2, 1fr); }
  .footer-grid { grid-template-columns: 1fr 1fr 1fr; }
  .article-layout { grid-template-columns: 1fr; }
  .article-sidebar { position: static; }
}

@media (max-width: 768px) {
  .hero-band h1 { font-size: 36px; letter-spacing: -0.72px; }
  .section-title { font-size: 28px; }
  .cards-grid-3, .cards-grid-2 { grid-template-columns: 1fr; }
  .footer-grid { grid-template-columns: 1fr 1fr; }
  .nav-links { display: none; flex-direction: column; position: absolute; top: 64px; left: 0; right: 0; background: var(--canvas); border-bottom: 1px solid var(--hairline); padding: 20px 24px; gap: 16px; }
  .nav-links.open { display: flex; }
  .nav-hamburger { display: flex; }
  .top-nav .container { position: relative; }
  .article-header h1 { font-size: 32px; letter-spacing: -0.5px; }
  .page-header h1 { font-size: 32px; }
  .contact-form { padding: 24px; }
}

@media (max-width: 640px) {
  .hero-band h1 { font-size: 32px; }
  .footer-grid { grid-template-columns: 1fr; }
  .footer-bottom { flex-direction: column; align-items: flex-start; }
  .cookie-banner .container { flex-direction: column; align-items: flex-start; }
}